Possible traces of feeding by beetles in coniferophyte wood from the Kazanian of the Kama River basin

Possible traces of feeding by woodboring beetle larvae (Coleoptera, ?Permocupedidae) are described from the Middle Permian locality Tikhie Gory (Tatarstan, right bank of the Kama River, Kazanian Stage). The traces in the form of perpendicular tunnels and shafts were left in picnoxylic wood of some conif erophytes s.l. (the orders Pinales, Cordaitales, or Vojnovskyales).