Mammal tracks record anthropogenic environmental influence in late-Holocene inland dunes of the European Sand Belt, Poland

Structures previously suggested to be footprints or tracks, but which have been little described, were observed within Quaternary inland dunes of the European Sand Belt. Excavation of these in several localities of the eastern part of the belt in Poland reveal that they are hoofprints of various ungulate mammals, both wild and domestic, as well as footprints of humans, ranging from potentially as early as the fifth to sixth century to the 19th century or later. Trample grounds and horizons with high densities of tracks are associated with other signs of intensifying usage of dune habitats for pasture and farming, including constrained movement of livestock, relatively well-developed paleosols, and evidence of ploughing. The presence of such abundant life traces not only reflects the presence of individual animals and people in an area, but sheds light on the potential processes and feedbacks that the tracemakers were involved in, in terms of maintaining and modifying the habitats themselves, which were dynamic and changeable. Footprints and their associated sedimentological records can be integrated with archaeological and historical data to analyse anthropogenic influence through time on physical landscapes and on the wild biota.