The trace fossil Circulichnis as a record of feeding exploration: New data from deep-sea Oligocene–Miocene deposits of northern Italy

Circulichnis is a puzzling, ring-like trace fossil preserved on bedding planes. It is represented mostly by its type ichnospecies C. montanus, which is characterized by an evenly circular or elliptical course. A new ichnospecies, C. ligusticus, is distinguished based on material from deep-sea Oligocene–Miocene deposits of the Tertiary Piemonte Basin, NW Italy. It shows a winding or irregular course. A new model of Circulichnis is proposed as an exploration burrow produced mostly by the trace makers of Helminthoidichnites, Gordia, or Helminthopsis, as it moved to a different sediment layer to checking its feeding utility. This type of behaviour occurred already by the Ediacaran and is common in marine, mostly deepsea bedded sediments. The trace makers belong mostly to polychaetes in marine sediments and to oligochaetes in continental sediments