Rediscovery of type material of the bioerosional trace fossil Talpina von Hagenow, 1840 and its ichnotaxonomic implications

Friedrich von Hagenow is renowned for his seminal inventory of the diverse invertebrate fauna from the Lower Maastrichtian chalk (Upper Cretaceous) of the Isle of Rügen, Germany. Tragically, his immense collection at Stettin (today Szczecin Poland) was almost completely destroyed during World War II, including the type material of hundreds of species established by him during the first half of the 19th century. Seven belemnite specimens from his collection rediscovered at the Museum fur Naturkunde Berlin, as well as 14 belemnites traced at the Museum fur Mineralogie und Geologie at the Senckenberg Naturhistorische Sammlungen Dresden, include syntypes of four Talpina ichnospecies, labelled T. ramosa, T. solitaria, T. foliacea and T. sentiformis. To document this material, a new ‘digital cylinder seal technique was applied. The rediscovered material allows lectotype designations for T. ramosa and T. solitaria (now Trypanites solitarius), whereas T. foliacea and T. sentiformis, which where not formally published by von Hagenow, are regarded as nomina nuda. As a consequence, the non topotypical neotypes of T. ramosa and T. solitaria, as designated by Ehrhard Voigt in 1972, are obsolete.