Ecostratigraphy, zonation and global correlation of earliest Ordovician planktic graptolites

The succession of earliest Ordovician planktic graptolites (graptoloids) in 11 key sequences around the world is reviewed and used to assess their distribution in depth facies, in paleolatitude and in time. An ecostratigraphic model shows the pattern of distribution of species across the shore-to-ocean profile in time. Fluctuations in this pattern are related to eustatic change; the Acerocare, Black Mountain, Ceratopyge and possibly, the Peltocare Regressive events are recognized. Surface-water temperature is found to have had little significant effect on distribution of Early Tremadoc species, and once the effects of facies is allowed for, their biostratigraphic utility can be assessed. A global suite of four chronozones is proposed for the Early Tremadoc and a further five zones comprise a provisional suite for the Late Tremadoc. The nine zones enable precise correlation of Tremadoc graptolitic sequences. The zones replace the previously used graptolite assemblages, are correlated with key conodont horizons and are, in upward sequence, the Zones of Rhabdinopora praeparabola, R. flabelliformis parabola, Anisograptus matanensis, R. f. anglica, Adelograptus, Paradelograptus antiquus, Kiaerograptus, Araneograptus murrayi-Araneograptus pulchellus, and Hunnegraptus copiosus