The halloporid trepostome bryozoans from the Ordovician Simpson Group of Oklahoma

The Bromide Formation of the Middle Ordovician Simpson Group of Oklahoma contains one of the oldest diverse bryozoan faunas in North America. The early divergence of many trepostome clades is revealed in these rocks. Three trepostome bryozoan species belonging to family Halloporidae are described from this fauna. Discriminant analysis is used to define the following halloporid species: Diplotrypa schindeli n. sp., Tarphophragma karklinsi n. sp., and Tarphophragma macrostoma (Loeblich). Preliminary cladistic analysis indicates that the family Halloporidae was already a distinct lineage by the Middle Ordovician. This suggests that by this time, many of the major trepostome clades were already established.