Emended Sandbian (Ordovician) conodont biostratigraphy in Baltoscandia and a new species of Amorphognathus

Conodonts are an important biostratigraphic tool for many Phanerozoic stages. Along with graptolites, they define all global Ordovician Stage boundaries. Within the Upper Ordovician interval, a known species of Amorphognathus tvaerensis (Bergström) is present in both Sandbian and Katian stratotype sections. Study of changes in the succession of A. tvaerensis revealed that elements in the upper part of its range differ morphologically quite distinctly from those in its lower part. Here, they are described as a new conodont species, A. viirae sp. nov. This new species is recognized in several Estonian and Swedish sections, with apparent occurrence also in Mójcza Quarry, Holy Cross Mountains, Poland and Black Knob Ridge, Oklahoma, USA. Detailed analysis of early Amorphognathus elements from Estonian and Swedish sections revealed the absence of A. inaequalis (Rhodes) in both regions, although a conodont subzone based on this species was identified earlier by some authors. Both the absence of A. inaequalis (Rhodes) and recognition of the new species A. viirae sp. nov. resulted in the revision of the conodont zonation, and a new version of it is proposed for the Sandbian Stage in the Atlantic Realm. The new zonation includes (from below) Pygodus anserinus, Baltoniodus variabilis, A. tvaerensis, B. gerdae, A. viirae and B. alobatus Conodont zones.