Immature Insect Assemblages from the Early Cretaceous (Purbeck/Wealden) of Southern England

The record of immature insects from the non-marine Purbeck and Wealden groups (Lower Cretaceous) of southern England is reviewed and expanded. Fossils of adult terrestrial insects are locally common, but terrestrial immature remains are restricted to transported hemipterans, most of which are sessile nymphs or puparia resembling those of extant whiteflies (Aleyrodidae). Remains of immature aquatic insects are more diverse and comprise the extant orders Plecoptera, Ephemeroptera, Odonata, Trichoptera, Hemiptera and Diptera. The Trichoptera are represented by larval cases constructed from a variety of materials corresponding to several ichnogenera. The Wealden immature insects were preserved in predominantly freshwater fluvial settings, whereas the Purbeck ones occur in lagoonal palaeoenvironments, ranging in salinity from brackish to hypersaline. The composition of aquatic immature insect faunas in the latter offers potential for palaeosalinity analysis, although there are complicating factors relating to habitat stability. Uncommon trace fossils such as beetle borings in wood provide evidence of immature insects not represented by body fossils.